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Anne Shirley is a freckle-faced, red-haired girl, who grows up in an orphanage having lost her parents at a very early age. Anne is always cheerful and fun-loving despite being brought up without love or affection. When she turns 11, she is adopted by the old farmer Matthew Cuthbert and his sister Marilla. Anne starts her new life with Matthew and Marilla at a farm called "Green Gables", but actually the Cuthberts wanted a boy who could help with their work on the farm... Overcoming many hardships and meeting many friends and people, Anne grows up to be a strong-minded woman.
Now in her twenties, Anne returns to Avonlea for the first time since Marilla Cuthbert's death. Gilbert has been offered a position in a hospital in New York, and he persuades Anne to come with him. He arranges a position for her at a large publishing house. Big city life isn't what they expected. Anne's manuscript is stolen by a dashing American writer, Jack Garrison. Thus the stage is set for a final three hour installment in the "Anne of Green Gables" story which follows the characters from New York, the war effort in Europe and eventually returns them to the red earth of Prince Edward Island.
At the turn of the century on Prince Edward Island, Matthew Cuthbert and his sister Marilla decide to take on an orphan boy as help for their farm. But they get an unexpected jolt when they're mistakenly sent a girl instead: Anne Shirley.
Anne Shirley, now a schoolteacher, has begun writing stories and collecting rejection slips. She acts as Diana's maid of honor, develops a relationship with Gilbert Blythe, and finds herself at Kingsport Ladies' College. But while Anne enjoys the battles and the friends she makes, she finds herself returning to Avonlea.
When Anne Shirley turns 13, she faces complex issues with her friends, inspirational adults and Marilla and Matthew. At the same time, she begins a friendship with Gilbert Blythe that emotionally escalates to disrupt the status quo of her peaceful world. Her free-spirited nature is challenged by her perceived need to become sensible, and her journey toward this goal is fraught with confusion and more than a few unfortunate – albeit, amusing - mishaps.
Anne heads to Charlottetown to attend accelerated classes as she continues to work towards her dream of being a school teacher. While there, she is forced to adapt quickly to her new surroundings and classmates as she navigates her way through her first school experience outside of her beloved Green Gables. Anne finds herself facing daunting choices for her future, the stirrings of romance, and tragedy unlike anything she's ever known. Meanwhile, Matthew and Marilla Cuthbert contend with advancing age as they deal with financial challenges and creeping ailments.